please find enclosed the final program of the EUROSILVA Workshop on
'Development and ageing in forest trees', to be held in Florence, Italy,
on 20-24 September, 2000.
The program can also be accessed on the workshop web page at:
Please circulate it as widely as possible.
Prof. Marco Borghetti
Prof Giuseppe Scarascia Mugnozza
Dr Federico Magnani
Workshop local organisers
DEVELOPMENT AND AGEING
IN FOREST TREES
20-24 September, 2000
Please visit the Workshop web page at:
What controls tree development and ageing? what is the relative role=20
of internal and external factors? At the tissue level, recent=20
advances have demonstrated in detail that development and senescence=20
both result from the interaction of genetic and environmental=20
controls, involving differential gene expression and activation. At=20
the tree level, a shift in hormonal patterns, possibly of endogenous=20
genetic origin, has been found to trigger the process of maturation,=20
which strongly affects the structure, function and growth of the=20
plant and initiates the diversion of resources from vegetative to=20
reproductive growth. Also at the tree and population level,=20
exogenous factors have been proposed to control the developmental=20
pattern of rise and decline of primary productivity with age. A=20
lively discussion has been going on for several years on what could=20
be the ecological determinants of the age-related decline in forest=20
productivity. Several theories have been proposed to explain this=20
phenomenon, variously suggesting nutrient or hydraulic limitations,=20
increased respiration and shifts in biomass allocation as possible=20
mechanisms, but to date there is no definitive evidence to support=20
any of them as a universal mechanism. Ageing and maturation are=20
generally thought to be intimately interwined albeith distinct=20
processes. Eventually, what are the implications of development and=20
ageing for the response of tissues and plants to their biotic and=20
Because of the variety of processes involved in tree development and=20
maturation, definite answers often result elusive, requiring a=20
comprehensive and holistic approach that takes both endogenous and=20
external factors into account.
The aim of the workshop is therefore to bring together experts of=20
different background and expertise to discuss these processes of=20
central relevance both to plant biology and to forest ecology moving=20
from different perspectives. An interdisciplinary workshop on this=20
issue that brings together foresters, ecologists and biologists seems=20
most appropriate and relevant to the aims of the EUROSILVA Action.
TREE PHYSIOLOGY SPECIAL ISSUE
Tree Physiology, a well-known refereed journal, has agreed to draw=20
from the Workshop a special issue on 'Development and ageing of=20
forest trees'. All the authors of accepted oral presentations are=20
invited to contribute, although only those papers that are deemed=20
acceptable by the reviewers and editor will be published in the=20
special issue of Tree Physiology.
Papers should be prepared in accordance with Tree Physiology's guide=20
for contributors and submitted (original plus three copies) to the=20
workshop local organisers no later than 31 October, 2000. The length=20
of the manuscript should not exceed 8 pages (about 8000 words,=20
including references). The manuscript should also be provided as a=20
digital file (a PDF file, if possible, otherwise an MS Word or Word=20
Perfect file, on an IBM-format floppy disk, zip disk or CD-ROM). The=20
file should be named after the corresponding author, e.g. smith.doc,=20
and contain all components of the article including tables, figures=20
and figure captions.
WORKSHOP FINAL PROGRAMME
Wednesday, 20 September
15.00 Arrival and registration
17.30 Opening session
18.30 Working group meeting
Thursday, 21 September - WG1 Growth and development
8.30 Valjakka M., Tuhkanen E., Vapaavuori E., H=84ggman H., Kangasjarvi J.
Keynote: Gene expression during leaf development and senescence=20
in birch (Betula pendula)
9.00 Rohde A., Boerjan W.
Keynote: Molecular components of terminal bud formation in poplar
9.30 Palva E.T., Heino P., Li C.Y., Puhakainen T., Welling A., Boije=20
M., Aalto O.
Development of dormancy and winter hardiess in birch
9.50 Collins A., Jones H., Campbell M.
The control of meristem establishment, maintenance and=20
maturation in Eucalyptus
10.40 Moritz T., Eriksson M., Israelsson M., Sandberg G., Olsson O.,=20
Olsen J., Junttila O.
Keynote: Studies of photoperiodic induction of shoot elongation=20
in trees: a transgenic approach
11.10 Magel E., Sundberg B., Uggla C.
Sucrose synthase is the dominating sucrose cleaving enzyme in=20
secondary differentiation processes of forest tree=20
axes, cambial growth and heartwood formation
11.30 Piispanen R., Saranpaa P.
Seasonal variation in storage lipids and non-structural=20
carbohydrates of young silver birch (Betula pendula=20
11.50 Taylor G., Robinson K.M., Ferris R., Bunn S.M., Bradshaw H.D.
Leaf growth and yield in poplar: can putative QTL be identified?
14.00 Poster session
15.00 Drouet A.
Keynote: Ageing control on flavonoid biosynthesis in walnut
15.30 Diaz-Sala C.
Keynote: Maturation and rejuvenation in trees: physiological,=20
molecular and cellular mechanisms
16.00 Barnett J.R., Eshtiaghi Z.
Morphological and anatomical rejuvenation induced by sequential=20
grafting in walnut
16.50 Fraga M.F., Centeno M.L., Ca=A4al M.J., Rodr=A1guez A., Fern ndez=20
B., Rodriguez R.
Molecular basis of organ and tree ageing and reinvigoration:=20
applications to micropropagation of selected=20
progenies and mature trees of Pinus radiata D. Don
17.10 Ritter E., Fernandez B., Rodriguez R., Giannino D., Racchi M.,=20
Wagner E., Albrechtowa J., Favre J.M., Smulders S.M.J.,=20
van der Linden G., Pasqualetto P.L., Paques M.
Development, validation and application of molecular,=20
morphological and physiological markers for juvenile=20
and mature state characterisation in woody plant=20
species (FAIR3- CT96-1445)
17.30 Danti S., Bagnoli F., Caparrini S., Racchi M.L.
Catalases are differentially expressed during development in=20
peach (Prunus persica)
17.50 Poster session
18.30 MC meeting
=46riday, 22 September - WG2 Mineral nutrition and water relations
8.30 Mencuccini M.
Keynote: Functional interpretation of allometric analyses in=20
forest trees. The role of hydraulic constraints
9.10 Kostner B., Falge E., Bernhofer Ch., Tenhunen J.
Age- and management-related effects on leaf area / sapwood area=20
relationships, canopy transpiration, and carbon gain of Picea abies=20
stands in Central Germany
9.30 Magnani F., Mencuccini M., Borghetti M.
Optimal self-similarity in xylem structure. Implications for=20
the developmental pattern of tree hydraulic architecture and=20
9.50 Hubbard R.M., Stiller V., Ryan M.G., Sperry J.S.
Stomatal conductance and photosynthesis vary linearly with=20
plant hydraulic conductance in ponderosa pine
10.40 Gower S.T.
Keynote: Towards a better understanding of age-related forest=20
11.20 Bond B.J., Ryan M.G., Phillips N., McDowell N.G.
Testing the hydraulic limitation hypothesis in tropical and=20
11.40 Nikinmaa E., M=84kel=84 A., Berninger F., Hari P., Peram=84ki M.,=
Size related decline of tree productivity from carbon balance=20
14.00 Poster session
14.40 Ryan M.G., Bond B.J., Hubbard R.M., Williams M.
Keynote: Experimental evidence for hydraulic constraints on=20
stomatal function, and implications for age-related decline in=20
15.20 Rust S., Roloff A.
Reduced photosynthesis in old oak (Quercus robur L.): the=20
impact of crown and hydraulic architecture
15.40 Niinemets U.
Changes in foliar morphology and chemical composition with=20
increasing tree size: a review of several case studies
16.00 Giardina C., Ryan M., Fownes J., Binkley D.
Total belowground carbon allocation and aboveground net primary=20
production in a fast growing Eucalyptus plantation
16.50 Urbinati C., Carrer M., Anfodillo T.
Age-dependent radial growth differences of Norway spruce (Picea=20
abies Karst.) in Eastern Italian Alps
17.10 Vanninen P.
Carbon budget for individual Scots pine trees: effects of size,=20
competition and site fertility on growth allocation
17.30 Bauer G.A., Bazzaz F.A.
Nitrogen partitioning influences photosynthetic capacity during=20
17.50 Poster session
18.30 MC meeting
Saturday, 23 September - WG3 Biotic and abiotic interactions
8.30 Ceulemans R., Bortier K., Gielen B., Laureysens I., Janssens I.A.
Keynote: Developmental processes in trees: effects of abiotic=20
factors, competition and genotype
9.10 Marek M.V., Sprtova. M.
Thinning effects on photosynthetic characteristic of a Norway=20
9.30 Stewart J.
Understanding the biological processes underlying growth=20
stagnation in lodgepole pine and its response to=20
9.50 Wirth C., Schulze E.-D.
Multiple control of variability of above-ground NPP in Siberian=20
Scots pine forests
10.40 Kuppers M.
Keynote: Ecophysiology of forest succession: from leaf to plant=20
in a competitive environment
11.20 Valentini R., Manca G., Dore S., Tedeschi V., Tirone G.
Age related carbon dynamics in a Quercus cerris L. coppice=20
forest under intensive management
11.40 Carrer M., Urbinati C., Anfodillo T.
Age influence in tree ring growth response to climate of=20
Larix decidua Mill. and Pinus cembra L. at timberline
14.00 Poster session
14.40 Kolb T.
Keynote: Ageing as an influence on tree response to ozone.=20
Theory and observations
15.20 Wieser G.
Can tree age dependent differences in ozone susceptibility of=20
Picea abies be related to cumulative ozone uptake and ozone flux ?=20
15.40 Havranek W.M., Wieser G., Tegischer K.
Ozone effects in a multiple stress experiment with young Norway=20
16.00 Vapaavuori E., Riikonen J., Oksanen E., Repo T., Peltonen P.,=20
Holopainen T., Holopainen J., Julkunen-Tiitto R.
Responses of fast-growing ozone-tolerant and ozone-sensitive=20
silver birch clones to elevated CO2 and O3: clonal differences and=20
results from first year of fumigation
16.50 Scarascia Mugnozza G., Calfapietra C., Sabatti M., de Angelis=20
P., Ceulemans R., Gielen B., Miglietta F.
Biomass growth and canopy development under elevated CO2=20
conditions of a poplar tree plantation: the POPFACE experiment
17.10 Ferris R., Sabatti M., Miglietta F., Mills R.F., Taylor G.
Leaf cell expansion and leaf cell production in poplar are both=20
increased in elevated CO2: a free-air CO2 enrichment (POPFACE)=20
17.30 Schutzendubel A., Langenfeld-Heyser R., Fritz E., Teichmann T.,=20
Ott, Godbold D., Polle A.
Cadmium-induced oxidative stress in ecomycorrhizal fungi and hosts
17.50 Poster session
18.30 MC meeting
Sunday, 24 September
9.00 Field excursion: Tuscanian forests and vineyards
18.00 Return to Florence
21 September - WG1 Growth and development
Abe H., Nakai T. Effects of the temporal deficit on morphogenesis of=20
the xylem cells in Cryptomeria japonica D. Don
Benderoth G., Silber G., Koloupaev V. Mechanical limits for the=20
growth of tree stems
Climent J., Chambel M.R., P=82rez E., Gil L. Prediction of heartwood=20
radius in Pinus canariensis Chr. Sm. Ex DC.
=46unada R., Iwatate-Suzuki T., Utsumi Y., Suzuki T., Sano Y. Cambial=20
reactivation in the
ring-porous hardwood Fraxinus mandshurica var. japonica
Giovannelli A., Giannini R. In vitro growth characteristics of mature=20
and re-invigorated chestnut (Castanea sativa Mill.)
Palumbo M.G., Giannini R., Paffetti D. Characterization of different=20
growth stages in Cupressus sempervirens L.
Kauppinen L., Immanen J., Ulvila J., Paulin L., Palva T., Helariutta=20
Y. Analysis of wood
development in birch by cDNA sequencing
Kurth W., Anzola J=81rgenson G., Dzierzon H., Schulte M. Digital=20
reconstruction of tree architecture as a tool for assessing growth=20
and functional performance
Pirttila A.M., Laukkanen H., Hohtola A. Differences in the=20
regeneration capacity of mature and juvenile Scots pine tissues
Saranpaa P., Piispanen R. Neutral and phospholipids in sapwood and=20
heartwood of Scots pine (Pinus sylvestris L.)
Schaber J., Badeck F. Physiology-based phenology models of forest trees
Tegischer K., Tausz M., Grill D., Wieser G. Tree-age and needle-age=20
dependent variations of antioxidants and photoprotective pigments=20
in spruce needles
Valjakka M., Luomala E.-M., Vapaavuori E., Sutinen S., Kangasjarvi=20
J., and Haggman H.
Photosynthesis, growth and senescence in sense-RbcS=20
transformed birch (Betula pendula) lines
Zaspel I., Hertel H. Development of progenies of Quercus petraea and=20
Q. robur descending from selected old trees of a relic population
22 September - WG2 Mineral nutrition and water relations
Anttonen S., Jolkkonen A., Linder S., Lundmark T., Vapaavuori E.=20
Effects of optimal fertilisation on carbon allocation, and=20
chemical composition of Norway spruce
Aranda X., De Herralde F., Fleck I., Sav=82 R. Hydraulic conductivity=20
in Quercus ilex resprouts after fire
Gall R., Landolt W., Bucher J.B. Are reversible changes in bark size=20
driven by the descent of assimilation products in Norway spruce=20
Genenger M., Brodbeck S., Zimmermann S., Frossard E., Brunner I.=20
Nitrate reductase activity of Norway spruce fine roots as=20
affectedby nitrogen and wood-ash fertilisation
Iivonen S., Rikala R., Vapaavuori E. Seasonal patterns of root and=20
shoot growth, gas exchange and carbohydrate status of Scots=20
pine seedlings subjected to low or high nutrient availability
Kutnar L., Levanic T. Growth and age characteristics of the Norway=20
spruce (Picea abies (L.) Karst.) from the mires on the Pokljuka=20
Landolt W., Michellod V., Bleuler P., Bucher J. Effects of soil=20
fertility, elevated CO2 and increased N deposition on=20
biochemical parameters in the foliage of young beech and spruce=20
Mencuccini M., Grace J. Age-related dynamics of carbon exchange in=20
European forests: the EU Carbo-Age Project
Oberhuber W. Vulnerability of an inner Alpine drought-exposed forest=20
ecosystem to climatic extremes
Patz G. The water balance in trees
Patakas A., Noitsakis B., Radoglou K., Jarvis P. Relationships=20
between photosynthetic rate and leaf anatomy in two evergreen oak=20
Radoglou K., Raftoyannis Y. Seasonal variation in physiological=20
parameters of broadleaved
seedlings during the first two years of field establishment
Whitehead D., Ryan M.G. Low leaf-specific hydraulic conductivity=20
reduces stomatal conductance and photosynthesis in older,=20
taller mountain beech trees
Zitnik S., Muller C., Cl=C7Cment A., Bonnet-Masimbert M., Hanke D.E.,=20
Kraigher H. Metabolism of sugars and phytic acid during long term=20
storage of acorns at low temperatures
23 September - WG3 Biotic and abiotic interactions
Einhorn K.S. Growth of beech (Fagus sylvatica) seedlings in response=20
to temperature at four different light levels in simulated=20
Gielen B., Calfapietra C., Ceulemans R. Effects of elevated CO2 on=20
crown structure, leaf area and growth of poplar genotypes in the=20
Kozovits A.R., Grams T.E.E., Blaschke H., Sommerkorn M., Matyssek R.=20
beech (Fagus sylvatica) and spruce (Picea abies) saplings=20
Muller M., Stabentheiner E., Tausz M., Wonisch A., Grill D.=20
Structural and physiological responses of forest tree species to=20
different environmental situations
Reiter I., Haberle K.-H., Blaschke H., Matyssek R. Quantifying=20
competition on crown level
between mature European beech and Norway spruce
Jokela A., Cordeiro A., Altabella T., Sarjala T., Bortolotti C.,=20
Tiburcio A., Huttunen S. Cloning and molecular analysis of=20
arginine decarboxylase (ADC) from Scots pine
La Mantia T., Cullotta S., Marchetti M., Barbera G. Ecophysiology of=20
holm oak (Quercus ilex L.) in different environmental=20
conditions in Sicily
Lomas C., Stober C., George E. Chronosequence studies of tree fine=20
root growth and turnover
Michelozzi M., Tognetti R., Rossi F. Seasonal variations of=20
photosynthetic capacities and total chlorophyll contents in Aleppo=20
pine trees from Italian provenances
Paoletti E., Raddi P., Di Lonardo V. Evolution of the Cupressus=20
sempervirens - Seiridium
cardinale interaction in elevated CO2 over time
Petkovsek S.A.S., Kraigher H. Impact of pollution on biodiversity of=20
types of ectomycorrhizae
Trost T., Gaberscik A. Responses of Norway spruce (Picea abies (L.)=20
Karst.) to enhanced UV-B radiation
Vilhar U., Rupel M., Diaci J., Kraigher H. Norway spruce regeneration=20
and interactions in the mycorrhizosphere
Please note: all forms and relevant information can be found on the=20
Workshop web page at:
Workshop Venue and Secretariat
All the working sessions will be held at the Educatorio di Fuligno -=20
Via Faenza, 48 - 50123 Florence (Italy) - ph. ++39 055 210232, a=20
former convent newly transformed into conference venue, within=20
walking distance of Florence city centre. The Secretariat will open=20
at 2:30 p.m. on Wednesday, 20 September and will remain open for the=20
duration of the Workshop. Accomodation will be in nearby hotels.=20
=46lorence, possibly the most charming of all cities, can be easily=20
reached either by plane or by train.
How to register
All participants must register by filling in the registration form=20
(available on the workshop web page) and returning it, with payment,=20
to the Organizing Secretariat Enic.
Registration Fees (VAT included)
By bank check/money order:
until 30 April 2000 - EUR 200
after 30 April 2000 - EUR 250
By credit card:
until 30 April 2000 - EUR 207
after 30 April 2000 - EUR 259
The fee includes: abstract book, congress kit, coffee breaks, working=20
lunches and the social dinner. Please remember that registration=20
forms received without payment will not be honored.
You may pay your congress fee by:
a) Bank check - payable to Enic/EUROSILVA
b) Money order - payment must be received by ENIC/EUROSILVA - account=20
number 23884/00 (ABI/CAB 06160/02805), Cassa di Risparmio di Firenze,=20
Agenzia n. 5, Via Gioberti n. 163/r, Florence, Italy. Payments are to=20
be made net of all bank charges which are to be paid by the=20
Note If you pay by bank transfer, please attach a copy of the=20
transfer order to your registration form.
c) Credit card - we accept VISA, CARTA SI, MASTERCARD and EUROCARD.=20
Please remember to fill in the pertinent sections of the registration=20
Refunds will be made after the closing of the workshop (upon written=20
request submitted to the Organizing Secretariat Enic via fax) as=20
before 30 June, 2000: 25% of the total amount will be withheld
after 30 June, 2000: no refunds will be made
A field excursion will be organized on Sunday 24 September, 2000.=20
Local foresters and ecologists will take us through the Chianti=20
region to the oak and beech stands of Montagnola Senese and of the=20
Parco Naturale della Val d'Orcia, close to the Monte Amiata massif.=20
We will then move on to visit the medieval abbey of Sant'Antimo,=20
founded by Charlemagne in 871, and the medieval village of=20
Montalcino, home to one of the most famous Italian wines, the=20
Brunello. A guided tour of the cellars in one of the most interesting=20
wineries in the area, complete with a wine tasting experience, will=20
end the day. Please note that an additional EUR 50 will be charged=20
for the field excursion.
The Organizing Secretariat Enic has reserved a sufficient number of=20
rooms in hotels located near the Workshop venue. Please fill in the=20
appropriate spaces on the registration form and return it to Enic.=20
Reservations cannot be accepted without a first night deposit + EUR=20
13 for booking fees. The deposit, minus the booking fees, will be=20
deducted from your hotel bill upon presentation of the voucher you=20
will receive from Enic. Please note that the hotel balance must be=20
paid in Italian Liras directly to the hotel upon departure.
Please specify your accomodation requirements in the registration=20
form (available in Acrobat .pdf or MS-Word .zip format) and return=20
it, along with the first night deposit for the category of your=20
choice, no later than June 30th. We cannot guarantee room=20
availability after that date since the number of rooms in downtown=20
=46lorence is limited.
Rooms will be assigned on a first-come-first-served basis. If the=20
single rooms are sold out, we will assign double rooms for single=20
occupancy; when rooms are no longer available in the category of your=20
choice, we will automatically book you into the next highest=20
category. Your consent will be asked before taking any actions.
Cancellations: written cancellations received before July 30thwill be=20
entitled to a refund of the first night deposit less EUR 26 for=20
administrative fees. No refunds will be made for cancellations=20
received after June 30th. All refunds will be made after the closing=20
of the workshop.
How to get to Florence
By plane: The Amerigo Vespucci Airport is just 3 km from the center=20
of Florence and offers service to Europe's main cities, as well as=20
the Rome and Milan airports with connections to all international=20
destinations. There is also a shuttle service with hourly departures=20
(price Lit. 6.000, tickets can be purchased at the airport espresso=20
bar or on board). Taxis are available on the "Arrivals" side of the=20
airport, the average fair to downtown Florence is Lit. 30.000. The=20
Pisa airport (80 km from Florence) is linked to Florence by trains=20
leaving every hour.
By train: Florence has excellent rail connections to the major=20
Italian and European cities via the Eurostar trains. The Santa Maria=20
Novella Railroad Station is in the center of the city.
By car - Parking: You can access the city by car although the=20
historic center is closed to private traffic. You can get to your=20
hotel by showing the booking voucher, and then you can park in the=20
hotel's nearby garages. The main underground public parking=20
facilities (hourly or daily rates) are located under the Santa Maria=20
Novella Railroad Station, under the Parterre in Piazza della Libert=85=20
and at the Fortezza da Basso.
Taxis: The minimum fare for a cab ride in Florence is Lit. 7.000.=20
The radio cab numbers are: 055 4798; 055 4242 and 055 4390. There are=20
cab stands at various strategic points throughout the city. You=20
cannot hail cabs on the street.
Local Scientific Organizer Dr. Federico Magnani CNR - IMGPF Via A.=20
Vannucci, 13 - 50134 Florence (Italy) Phone ++39 055 461453 - Fax=20
++39 055 486604 E mail: federico at imgpf.fi.cnr.it
Organizing Secretariat Enic Viale Amendola, 20 - 50121 Florence=20
(Italy) Phone ++39 055 240275 - Fax ++39 055 2345078 E mail:=20
chiara at egr.it- Website: http://www.egr.it
Prof. Satu Huttunen, Finland
Dr. Jurg Bucher, Switzerland
Prof. Bjorn Sundberg, Sweden
Prof. Paul G. Jarvis, United Kingdom
Prof. Rainer Matyssek, Germany
Prof. Giuseppe Scarascia Mugnozza , Italy
Prof. Marco Borghetti , Italy
Dr. Federico Magnani, Italy
CNR - National Research Council, Italy
SISEF - Italian Society of Silviculture and Forest Ecology
Italian Academy of Forest Science